Wednesday, August 21, 2013

THE LABRADOR RETRIEVER GENOTYPE'S AND THE BREED STANDARD

 This is a guest post by our friend Michael Marshburn of Jam Up Retriever's



 For the sake of this article genotype's expressed are as follow's

B = Black and is dominant
b = Brown and is recessive.
E = Color approval and is dominant
e = Color cancel and is recessive.

All source's quoted in this article are referenced
bellow!


Michael is a young, avid trainer/breeder. He currently trains his own personal dogs for hunting and a few for retriever hunt tests. Michael believes that it is important to strive for healthy dogs that can perform for their intended purpose and to spread the word to potential buyers about qualities they should seek in their prospected hunting partners and pets. When the calls are hung up and the dogs are through with their work, Michael can be found on the job at his fire department or assisting his bride-to-be with her FFA chapter activities.


Having been the most registered dog for the last thirty, or so, years, it’s an understatement to say that the Labrador Retriever is in demand. With this demand comes the demand for more people able to produce puppies, which isn’t always in the best interests of the breed. The parent club, the Labrador Retriever Club of America, has set the breed standard for the dog that it represents. The LRC is the body that is the member of the AKC, which allows for the registration of labs with their registry. This standard is the basis for the conformation events, which are a visual judging of the animal’s conformance to the standards the club has set.

            Everyone has always heard how field trial labs look so different from show labs, although it hasn’t always been this way. 3xNFC CFC DCh. Shed of Arden was a very accomplished specimen of the breed. He demonstrated what we all should strive for: a strong visual representation of the standard that performs the initial intent of the breed. However, field trials were a different ball game back in 1946. As a matter of fact, Shed won the 1946 National Open after completing a double water blind of fifty yards, according to Hunting Lab Pedigree. We have changed the game of trials over the years and have required more of these wonderful animals, and have thus caused their physique to change. The largest change you will see comes from a more slim face and a chest that isn’t as deep.

            But, the labs of today are beginning to change even more. One look at any classified and you will begin to notice a spectrum of colors that are beginning to emerge. Silver, charcoal, champagne and white are the flavors of this week. What does all of this mean? Well, let us first identify what the breed standard has to say about the coat color. As direct from the LRC website, “The Labrador Retriever coat colors are black, yellow and chocolate. Any other color or a combination of colors is a disqualification. A small white spot on the chest is permissible, but not desirable. White hairs from aging or scarring are not to be misinterpreted as brindling. Black - Blacks are all black. A black with brindle markings or a black with tan markings is a disqualification. Yellow - Yellows may range in color from fox-red to light cream, with variations in shading on the ears, back, and underparts of the dog. Chocolate - Chocolates can vary in shade from light to dark chocolate. Chocolate with brindle or tan markings is a disqualification.”

            To begin with, let’s look at the basic color that labs originated from: black. Labs originated from the St. John’s Water Dog, which were known to be black with white patches on their chests. This is why the white spot is not considered a mismark, but preference is given to ones with smaller markings. Yellows have the only variety for color, of which the first yellow, Ben of Hyde, was stated to have been butterscotch. So, the first yellow was of the “fox red” shade, and more labs were reported to have been called “gold” until the ruling came that yellow described the color of the dog and that “gold” describes an element. The “light cream” is what many people are now describing as “white.” Chocolate, in and of itself, is self-explanatory, much like the black.

            So, now that the colors are established, I will begin to guide you into the “genotypes” of these dogs. I promise, this is part of a grand scheme to help you better understand how these designer colors play in. As always in genetics, each offspring receives two copies of a gene; One from each parent. A labs coat color is determined from two genes, E and B. Through use of the Punnett’s Square, you can determine possible litter outcomes based on the two dogs genotypes.



A basic black labs genotype is described as EEBB. When a basic black lab is bred to another Basic black lab, both parents only have the option of giving dominant genes. Therefore, all offspring from them will be black with the genotype EEBB. A yellow labs genotype is described as eeBB. When two yellows with no recessive genes are bred, they will always produce eeBB puppies. A chocolate is described as EEbb, therefore, as with the previous scenarios, they will produce all chocolate puppies when bred to another chocolate of the same genotype.

            The next part of this may seem to get a little tricky, but, I promise, it isn’t hard to keep up with. Let’s go back to the black genotype. Now, some of you are probably very curious as to how blacks can sometimes throw yellows, chocolates or both. To begin with this area, you have to understand that dominant genes are genes that show, or become the dog’s phenotype. A black lab can carry yellow or chocolate or both, depending upon which genes its parents gave during conception. So, a black lab carrying yellow would be EeBB and one carrying chocolate would be EEBb. One carrying both would be EeBb. So, basically, we’ve established that blacks can carry both yellow and chocolate and that when bred to another dog that either displays, or carries recessively, these traits, they can produce the different varieties of colors. So, can yellows or chocolates carry recessives? Absolutely. Yellows can carry recessive chocolate (eeBb), only and chocolates can carry recessive yellow (Eebb), only. If you’ve caught on to all of this, you can feel relieved that you don’t have to remember the technical details and someone else has a resource you can use to determine possible litter outcomes. Blue Knight Labradors has a Coat Color Inheritance chart that has every genotype matching along with litter outcome potential.

            So, if you’re inquisitive, you’ve realized that there is one more genotype that I didn’t discuss. The “Dudley,” or eebb genotype, is a dog lacking any black pigment. They are described as having no pigment around the nail beds or eyes. The Dudley is a disqualification in the show ring and is considered a mismark puppy. They are also currently being purported as “champagne” labs and are being sold largely on the grounds of being a “rare” coat color. Quite simply, the chance of producing them occurs any time breeding pairs contain any of the combinations where yellow and chocolate are present together. What does this mean? It means your dog will be disqualified in the show ring. It means absolutely nothing along the lines of how well the dogs will retrieve or perform.

 





So, where do the rest of these colors play in? The other two colors fall into a category of an outlying gene that is in debate of its place in the Labrador world. The LRC’s official stance is that the only accepted coat colors of black and chocolates involve no description of shading. This dilute gene, commonly referred to as the recessive d gene, plays a part in the black and chocolate phenotypes causing a dilution of the pigment present and presenting the described “charcoal” and “silver” labs. These coat colors have been mandated by the AKC to register as Black and Chocolate in color, respectively. Basically, yet again, even though these described coat colors are considered mismarks and disqualifications, it does not predict hunting ability or temperament.

            The main argument that is presented against these coat colors is that, according to the LRC, there is plenty of evidence present to conclude that the recessive dilute gene was not present in the Labrador at the time of its founding. The LRC also makes the statement that the only known breed that the dilute gene makes a characteristic which is standard is the weimaraner. They also state that the argument of the dilute gene being a “shade” of the black and chocolate coat colors is inaccurate and that the causes for shading differences in yellows does not rely on the dilute gene.

            By and large, the main argument that you will find on many silver breeders websites is that the AKC has made the statement that the dogs are registered and do appear to be purebred Labradors. However, one argument purported that is of highly inaccurate terms is that DNA tests were performed and it was proven that silvers were purebred Labradors. To begin with, all canines we know today are descendants of wolves. In performing a DNA test, one can only derive parentage and species of the dogs as well as known genetic anomalies. All you will ever find is that dogs have the same numbers of chromosomes and that they are all descendants of wolves. Dogs do not have any different numbers of genes based on breed, hence why a Chihuahua can impregnate a Great Dane (by principle, of course). One can’t look at a DNA map and specifically identify breed specifics of Shih Tzu or Terrier apart from a mutt.

            What is the significance of all of this? Well, first of all, it is to say that nothing other than a dog’s natural instinct given to it by the parents along with your training will ever determine an outcome. Also, by saying that, dogs with a history of trainability, shown by titles in a pedigree, demonstrate that throughout the dog’s ancestry their predecessors have conformed to a standard and achieved a title. More titles will help you determine if a dog has potential to learn the skill set you wish to instill upon them. However, one must keep in mind that not every Stanley Cup Champion has a child who achieves the same goals. By and large, the goal should be to obtain a healthy puppy who can be taught the skills which serve your needs best. This can be done by seeking out breeders who have breeding stock that possess the necessary health clearances and who show you performance pedigrees that demonstrate a history of dogs that achieved standards and earned titles along with a history of producing puppies that can achieve standards. Seeking these things will get you much further than seeking out the services from someone who doesn’t find these qualities as important.

 


 

Sources:


Labrador Retriever Club of America http://www.thelabradorclub.com/



 
 

 
 
 

 

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